Symptoms are produced on almost all aerial parts of the wheat plant but are most common on stem, leaf sheaths and upper and lower leaf surfaces. Uredial pustules (or sori) are oval to spindle shaped and dark reddish brown (rust) in color. They erupt through the epidermis of the host and are surrounded by tattered host tissue. The pustules are dusty in appearance due to the vast number of spores produced. Spores are readily released when touched.